Fossils
Fossils of Pakistan
Pakistan’s strata document the rise of apes and the birth of whales, alongside a billion years of marine life. Start with the landmark finds, or browse every fossil-bearing unit by group. 121 of 285 formations carry a recorded fossil note.
Landmark finds
Cenozoic
The first apes — Sivapithecus
The Potwar Siwaliks yielded Sivapithecus, an early great ape central to the debate over the origin of the orangutan lineage.
Cenozoic
The first whales
Eocene rocks of Punjab and Balochistan captured the move of mammals back to the sea — walking whales like Pakicetus and Ambulocetus.
Cenozoic
The Siwalik mammal fauna
River sediments shed off the rising Himalaya preserve one of Earth’s richest fossil-mammal records — elephants, rhinos, giraffes, horses, Deinotherium and Stegodon.
Mesozoic
Balochistan dinosaurs
Latest-Cretaceous beds of the Sulaiman region have produced titanosaur sauropod dinosaurs — among the few dinosaur finds in Pakistan.
Cenozoic
Eocene Nummulites
Coin-shaped larger foraminifera — Nummulites, Assilina, Alveolina — are so abundant they build entire limestones across the Eocene of Pakistan.
Precambrian
The earliest life — stromatolites
Layered microbial structures (stromatolites) and algal mats in the Precambrian record the oldest traces of life preserved in Pakistan’s rocks.
Browse by fossil group
Mammals11 Reptiles, fish & other vertebrates9 Foraminifera56 Molluscs43 Ammonites & cephalopods18 Corals18 Brachiopods22 Echinoderms15 Bryozoans9 Trilobites5 Plants10 Stromatolites & algae9 Trace fossils9 Conodonts & ostracods11
Mammals
11Chinji Formation
Neogene DeinotheriumHipparionSivapithecusCrocodilesTurtles
Chitarwata Formation
Paleogene Mammal fossilsPlant fossils
Drug Formation
Paleogene Walking whales (Ambulocetus)BasilosauridsLarger foraminifera
Kamlial Formation
Neogene Mammal fossilsFreshwater gastropods
Kuldana Formation
Paleogene PakicetusGandakasiaIchthyolestes
Lei Conglomerate
Quaternary Mammal fossilsStegodonEarly Paleolithic artifacts
Litra Formation
Neogene Middle Siwalik mammals (incl. hipparionine horses)
Manchhar Formation
Neogene Mammal fossilsPlant fossils
Murree Formation
Neogene Mammal bonesFreshwater molluscsPlant fossils
Nagri Formation
Neogene Mammal fossilsCrocodilian remainsFreshwater fauna
Vihowa Formation
Neogene Lower Siwalik mammals (e.g. Eotragus)
Reptiles, fish & other vertebrates
9Chinji Formation
Neogene DeinotheriumHipparionSivapithecusCrocodilesTurtles
Gaj Formation
Neogene ForaminiferaMolluscsMarine vertebrates
Khadro Formation
Paleogene oystersreptile bonesforaminifera
Mianwali Formation
Triassic Claraia bivalvesOphiceras ammonoidsConodontsFish remains
Nagri Formation
Neogene Mammal fossilsCrocodilian remainsFreshwater fauna
Nari Formation
Paleogene ForaminiferaMolluscsShark teeth
Rawat formation
Permian Permian invertebrates
Tredian Formation
Triassic Plant fragmentsVertebrate bones (rare)
Vitakri Formation
Cretaceous titanosaurian sauropodstheropod dinosaursmesoeucrocodylianspterosaurs
Foraminifera
56Aghil Limestone
Triassic–Jurassic ForaminiferaAlgae
Amb Formation
Permian Brachiopods (Productus)CrinoidsBryozoansForaminifera
Ashtigar formation
Jurassic Trigonia-type bivalvesbrachiopodscrinoidsforaminiferanannofossils
Borom Formation
Triassic ConodontsForaminifera
Darkot Group
Carboniferous–Permian bryozoansforaminiferabrachiopodscrinoidscoralsgastropods
Domanda Formation
Paleogene Nummulites beaumontiNummulites subbeaumontiAlveolina ellipticaDictyoconoides cooki
Drazinda Formation
Paleogene foraminiferabivalvesbryozoansechinoids
Drug Formation
Paleogene Walking whales (Ambulocetus)BasilosauridsLarger foraminifera
Dungan Formation
Paleogene larger foraminiferacorals
Gaj Formation
Neogene ForaminiferaMolluscsMarine vertebrates
Goru Formation
Cretaceous ForaminiferaAmmonitesBivalves
Gurguri Formation
Paleogene ForaminiferaMollusks
Habib Rahi Formation
Paleogene Larger foraminifera (Nummulites, Assilina)GastropodsEchinoids
Hangu Formation
Paleogene Miscellanea miscellaLockhartia haimeiLockhartia conditiLepidocyclina punjabensisOperculina canaliferaDaviesina longhami
Hoshab Formation
Paleogene–Neogene Planktonic foraminiferaTrace fossils
Khadro Formation
Paleogene oystersreptile bonesforaminifera
Khyber limestone
Carboniferous–Permian brachiopodsgastropodspelecypodsammonoidsbryozoansfusulinids
Kingriali Formation
Triassic Megalodont bivalvesGastropodsForaminifera
Kirthar Formation
Paleogene nummuliteslarger foraminifera
Lakhra Formation
Paleogene foraminiferacoralsmolluscs
Laki Formation
Paleogene foraminiferagastropodsbivalvesechinoidsalgae
Lockhart Limestone
Paleogene Operculina subsalsaMiscellanea miscellaMiscellanea stampiLockhartia haimeiLockhartia newboldiLepidocyclina punjabensisDiscocyclina ranikotensis
Loralai Formation
Jurassic AmmonitesBelemnitesBivalvesForaminifera
Margala Hill Limestone
Paleogene Nummulites mamillatusNummulites atacicusAssilina spinosaAssilina laminosaAssilina granulosaAssilina exponensLockhartia tipperiLockhartia conditiAlveolinaOperculinaMiliolidsGastropodsEchinoids
Mughal Kot Formation
Cretaceous AmmonitesForaminiferaBivalves
Nammal Formation
Paleogene Planktonic foraminiferaNannofossils
Nari Formation
Paleogene ForaminiferaMolluscsShark teeth
Nisai Limestone
Cretaceous–Paleogene Miscellanea miscellaOther larger foraminiferaAlgae
Ormara Formation
Neogene MollusksEchinoidsForaminifera
Panjgur Formation
Paleogene–Neogene ForaminiferaTrace fossils
Panoba Formation
Paleogene Globorotalia aequaGloborotalia quadrataAssilina pustulosaOrbitolites complanatusNummulites sp.
Parh Group
Cretaceous Planktic foraminiferaAmmonites
Parh Limestone
Cretaceous planktonic foraminifera (Globotruncana spp.)
Parkini Formation
Neogene Benthic foraminiferaMollusks
Patala Formation
Paleogene Planktonic foraminiferaCalcareous nannofossilsDinoflagellates
Pirkoh Formation
Paleogene foraminiferagastropodsbivalvesechinoids
Ranikot Formation
Paleogene Larger foraminiferaMollusksPlant fossils
Ranikot Group
Paleogene Larger foraminiferaGastropodsPlant fossils in upper part
Reshit Formation
Jurassic foraminiferaostracodscrinoidscoralsechinoidsbrachiopodsammonites
Reshun Formation
Cretaceous MegalodonCoralsForaminifera
Saindak Formation
Paleogene AssilinaFasciolitesEchinolampasGastropods (Velates)
Sakesar Formation
Paleogene Nummulites mamillatusNummulites atacicusAssilina spinosaAssilina granulosaAssilina laminosaAlveolinaLockhartia tipperiLockhartia conditiDiscocyclinaGastropodsEchinoids
Samana Suk Formation
Jurassic CoralsBrachiopodsEchinoidsForaminiferaAlgae
Sangiali Formation
Paleogene Larger foraminiferaGastropods
Sembar Formation
Cretaceous AmmonitesBelemnitesForaminifera
Shaheed Ghat Formation
Paleogene nummulitesgastropodsbivalves
Shaksgam formation
Permian Parafusulina sp.Paraschwagerina sp.Neoschwagerina sp.
Shekhan Formation
Paleogene Orbitolites complanatusAlveolina oblongaAlveolina vredenburgiAssilina daviesiAssilina laminosaAssilina dandoticaNummulites atacicusCoralsGastropodsEchinoids
Shinawari Formation
Jurassic AmmonitesBivalvesForaminifera
Siahan Formation
Paleogene Deep water foraminiferaTrace fossils
Sinjrani Volcanic Group
Cretaceous AlgaeForaminiferaCorals (in limestone interbeds)
Tekri formation
Jurassic spongesgastropodsostreidsforaminifera
Wargal Limestone
Permian Productid brachiopodsFusulinid foraminiferaCoralsGastropodsCephalopods
Yashkuk formation
Jurassic benthic foraminiferacalcareous algaenannofossilsostracods
Yasin group
Cretaceous Orbitolinarudists (Hippurites)
Zaluch Group
Permian brachiopods (productids)fusulinidsbryozoa
Molluscs
43Ashtigar formation
Jurassic Trigonia-type bivalvesbrachiopodscrinoidsforaminiferanannofossils
Darkot Group
Carboniferous–Permian bryozoansforaminiferabrachiopodscrinoidscoralsgastropods
Datta Formation
Jurassic Plant fossilsCoal tracesFreshwater bivalves
Drazinda Formation
Paleogene foraminiferabivalvesbryozoansechinoids
Gaj Formation
Neogene ForaminiferaMolluscsMarine vertebrates
Goru Formation
Cretaceous ForaminiferaAmmonitesBivalves
Gurguri Formation
Paleogene ForaminiferaMollusks
Habib Rahi Formation
Paleogene Larger foraminifera (Nummulites, Assilina)GastropodsEchinoids
Humai Formation
Cretaceous Hippurites (rudist bivalves)OrbitoidesLepidorbitoidesGastropods
Jiwani Formation
Quaternary MollusksCorals
Jutana Formation
Cambrian Trilobites (Redlichia noetlingi)Brachiopods (Lingulella fuchsia, Botsfordia granulata)Gastropods
Kamlial Formation
Neogene Mammal fossilsFreshwater gastropods
Khadro Formation
Paleogene oystersreptile bonesforaminifera
Khyber limestone
Carboniferous–Permian brachiopodsgastropodspelecypodsammonoidsbryozoansfusulinids
Kingriali Formation
Triassic Megalodont bivalvesGastropodsForaminifera
Lakhra Formation
Paleogene foraminiferacoralsmolluscs
Laki Formation
Paleogene foraminiferagastropodsbivalvesechinoidsalgae
Loralai Formation
Jurassic AmmonitesBelemnitesBivalvesForaminifera
Lun Shale
Triassic Ammonoids (rare)Bivalves
Margala Hill Limestone
Paleogene Nummulites mamillatusNummulites atacicusAssilina spinosaAssilina laminosaAssilina granulosaAssilina exponensLockhartia tipperiLockhartia conditiAlveolinaOperculinaMiliolidsGastropodsEchinoids
Mianwali Formation
Triassic Claraia bivalvesOphiceras ammonoidsConodontsFish remains
Mughal Kot Formation
Cretaceous AmmonitesForaminiferaBivalves
Murree Formation
Neogene Mammal bonesFreshwater molluscsPlant fossils
Musakhel group
Triassic ammonoids (Ceratites)bivalves
Nari Formation
Paleogene ForaminiferaMolluscsShark teeth
Nowshera Formation
Devonian coralsstromatoporoidscephalopodsgastropodsbrachiopodsconodonts
Ormara Formation
Neogene MollusksEchinoidsForaminifera
Pab Sandstone
Cretaceous BivalvesGastropodsTrace fossils
Panjpir Formation
Ordovician–Silurian crinoidspelecypodscephalopodsnautiloidsconodonts
Parkini Formation
Neogene Benthic foraminiferaMollusks
Pirkoh Formation
Paleogene foraminiferagastropodsbivalvesechinoids
Ranikot Formation
Paleogene Larger foraminiferaMollusksPlant fossils
Ranikot Group
Paleogene Larger foraminiferaGastropodsPlant fossils in upper part
Saindak Formation
Paleogene AssilinaFasciolitesEchinolampasGastropods (Velates)
Sakesar Formation
Paleogene Nummulites mamillatusNummulites atacicusAssilina spinosaAssilina granulosaAssilina laminosaAlveolinaLockhartia tipperiLockhartia conditiDiscocyclinaGastropodsEchinoids
Sangiali Formation
Paleogene Larger foraminiferaGastropods
Shaheed Ghat Formation
Paleogene nummulitesgastropodsbivalves
Shekhan Formation
Paleogene Orbitolites complanatusAlveolina oblongaAlveolina vredenburgiAssilina daviesiAssilina laminosaAssilina dandoticaNummulites atacicusCoralsGastropodsEchinoids
Shinawari Formation
Jurassic AmmonitesBivalvesForaminifera
Surghar Group
Jurassic ammonites (Bouleiceras)brachiopodsbivalves
Tekri formation
Jurassic spongesgastropodsostreidsforaminifera
Toi Formation
Paleogene freshwater bivalvesgastropodsplant remains
Wargal Limestone
Permian Productid brachiopodsFusulinid foraminiferaCoralsGastropodsCephalopods
Ammonites & cephalopods
18Bdongo-La formation
Jurassic coralsammonites
Chhidru Formation
Permian Last Paleozoic brachiopodsAmmonoidsDiminished fauna near top
Chidru Formation
Permian Last Paleozoic brachiopodsAmmonoidsDiminished fauna near top
Goru Formation
Cretaceous ForaminiferaAmmonitesBivalves
Khyber limestone
Carboniferous–Permian brachiopodsgastropodspelecypodsammonoidsbryozoansfusulinids
Loralai Formation
Jurassic AmmonitesBelemnitesBivalvesForaminifera
Lun Shale
Triassic Ammonoids (rare)Bivalves
Mianwali Formation
Triassic Claraia bivalvesOphiceras ammonoidsConodontsFish remains
Mughal Kot Formation
Cretaceous AmmonitesForaminiferaBivalves
Musakhel group
Triassic ammonoids (Ceratites)bivalves
Nowshera Formation
Devonian coralsstromatoporoidscephalopodsgastropodsbrachiopodsconodonts
Panjpir Formation
Ordovician–Silurian crinoidspelecypodscephalopodsnautiloidsconodonts
Parh Group
Cretaceous Planktic foraminiferaAmmonites
Reshit Formation
Jurassic foraminiferaostracodscrinoidscoralsechinoidsbrachiopodsammonites
Sembar Formation
Cretaceous AmmonitesBelemnitesForaminifera
Shinawari Formation
Jurassic AmmonitesBivalvesForaminifera
Surghar Group
Jurassic ammonites (Bouleiceras)brachiopodsbivalves
Wargal Limestone
Permian Productid brachiopodsFusulinid foraminiferaCoralsGastropodsCephalopods
Corals
18Ali Masjid formation
Devonian brachiopodscorals
Bdongo-La formation
Jurassic coralsammonites
Chalt formation
Carboniferous–Permian bryozoansFenestellaproductidscorals
Chiltan Limestone
Jurassic corals
Darkot Group
Carboniferous–Permian bryozoansforaminiferabrachiopodscrinoidscoralsgastropods
Dungan Formation
Paleogene larger foraminiferacorals
Jiwani Formation
Quaternary MollusksCorals
Jobra formation
Ordovician Cruziana rugosa
Lakhra Formation
Paleogene foraminiferacoralsmolluscs
Nowshera Formation
Devonian coralsstromatoporoidscephalopodsgastropodsbrachiopodsconodonts
Pir Sabak formation
Devonian corals
Reshit Formation
Jurassic foraminiferaostracodscrinoidscoralsechinoidsbrachiopodsammonites
Reshun Formation
Cretaceous MegalodonCoralsForaminifera
Samana Suk Formation
Jurassic CoralsBrachiopodsEchinoidsForaminiferaAlgae
Shekhan Formation
Paleogene Orbitolites complanatusAlveolina oblongaAlveolina vredenburgiAssilina daviesiAssilina laminosaAssilina dandoticaNummulites atacicusCoralsGastropodsEchinoids
Shogram Formation
Devonian rugose coralsbrachiopodsconodonts
Sinjrani Volcanic Group
Cretaceous AlgaeForaminiferaCorals (in limestone interbeds)
Wargal Limestone
Permian Productid brachiopodsFusulinid foraminiferaCoralsGastropodsCephalopods
Brachiopods
22Ali Masjid formation
Devonian brachiopodscorals
Amb Formation
Permian Brachiopods (Productus)CrinoidsBryozoansForaminifera
Ashtigar formation
Jurassic Trigonia-type bivalvesbrachiopodscrinoidsforaminiferanannofossils
Baroghil group
Ordovician–Silurian acritarchsconodontsbrachiopods
Chalt formation
Carboniferous–Permian bryozoansFenestellaproductidscorals
Chhidru Formation
Permian Last Paleozoic brachiopodsAmmonoidsDiminished fauna near top
Chidru Formation
Permian Last Paleozoic brachiopodsAmmonoidsDiminished fauna near top
Dandot Formation
Permian BrachiopodsBryozoans
Darkot Group
Carboniferous–Permian bryozoansforaminiferabrachiopodscrinoidscoralsgastropods
Gircha formation
Permian Trigonotreta stokesiSpirelytha petaliformisPunctospirifer afghanusEurydesma sp.
Jutana Formation
Cambrian Trilobites (Redlichia noetlingi)Brachiopods (Lingulella fuchsia, Botsfordia granulata)Gastropods
Khyber limestone
Carboniferous–Permian brachiopodsgastropodspelecypodsammonoidsbryozoansfusulinids
Kussak Formation
Cambrian Trilobites (Redlichia noetlingi)Brachiopods (Neobolus warthi, Lingulella)Hyolithes wynnei
Nowshera Formation
Devonian coralsstromatoporoidscephalopodsgastropodsbrachiopodsconodonts
Reshit Formation
Jurassic foraminiferaostracodscrinoidscoralsechinoidsbrachiopodsammonites
Samana Suk Formation
Jurassic CoralsBrachiopodsEchinoidsForaminiferaAlgae
Sardhi Formation
Permian BrachiopodsBryozoansCrinoids
Shogram Formation
Devonian rugose coralsbrachiopodsconodonts
Surghar Group
Jurassic ammonites (Bouleiceras)brachiopodsbivalves
Warcha Formation
Permian BrachiopodsBryozoans
Wargal Limestone
Permian Productid brachiopodsFusulinid foraminiferaCoralsGastropodsCephalopods
Zaluch Group
Permian brachiopods (productids)fusulinidsbryozoa
Echinoderms
15Amb Formation
Permian Brachiopods (Productus)CrinoidsBryozoansForaminifera
Ashtigar formation
Jurassic Trigonia-type bivalvesbrachiopodscrinoidsforaminiferanannofossils
Darkot Group
Carboniferous–Permian bryozoansforaminiferabrachiopodscrinoidscoralsgastropods
Drazinda Formation
Paleogene foraminiferabivalvesbryozoansechinoids
Habib Rahi Formation
Paleogene Larger foraminifera (Nummulites, Assilina)GastropodsEchinoids
Laki Formation
Paleogene foraminiferagastropodsbivalvesechinoidsalgae
Margala Hill Limestone
Paleogene Nummulites mamillatusNummulites atacicusAssilina spinosaAssilina laminosaAssilina granulosaAssilina exponensLockhartia tipperiLockhartia conditiAlveolinaOperculinaMiliolidsGastropodsEchinoids
Ormara Formation
Neogene MollusksEchinoidsForaminifera
Panjpir Formation
Ordovician–Silurian crinoidspelecypodscephalopodsnautiloidsconodonts
Pirkoh Formation
Paleogene foraminiferagastropodsbivalvesechinoids
Reshit Formation
Jurassic foraminiferaostracodscrinoidscoralsechinoidsbrachiopodsammonites
Sakesar Formation
Paleogene Nummulites mamillatusNummulites atacicusAssilina spinosaAssilina granulosaAssilina laminosaAlveolinaLockhartia tipperiLockhartia conditiDiscocyclinaGastropodsEchinoids
Samana Suk Formation
Jurassic CoralsBrachiopodsEchinoidsForaminiferaAlgae
Sardhi Formation
Permian BrachiopodsBryozoansCrinoids
Shekhan Formation
Paleogene Orbitolites complanatusAlveolina oblongaAlveolina vredenburgiAssilina daviesiAssilina laminosaAssilina dandoticaNummulites atacicusCoralsGastropodsEchinoids
Bryozoans
9Amb Formation
Permian Brachiopods (Productus)CrinoidsBryozoansForaminifera
Chalt formation
Carboniferous–Permian bryozoansFenestellaproductidscorals
Dandot Formation
Permian BrachiopodsBryozoans
Darkot Group
Carboniferous–Permian bryozoansforaminiferabrachiopodscrinoidscoralsgastropods
Drazinda Formation
Paleogene foraminiferabivalvesbryozoansechinoids
Khyber limestone
Carboniferous–Permian brachiopodsgastropodspelecypodsammonoidsbryozoansfusulinids
Sardhi Formation
Permian BrachiopodsBryozoansCrinoids
Warcha Formation
Permian BrachiopodsBryozoans
Zaluch Group
Permian brachiopods (productids)fusulinidsbryozoa
Trilobites
5Baghanwala Formation
Cambrian Trace fossilsTrilobites (rare)
Jobra formation
Ordovician Cruziana rugosa
Jutana Formation
Cambrian Trilobites (Redlichia noetlingi)Brachiopods (Lingulella fuchsia, Botsfordia granulata)Gastropods
Khewra Sandstone
Cambrian Trilobite trails (trace fossils)Skolithos burrowsCruziana traces
Kussak Formation
Cambrian Trilobites (Redlichia noetlingi)Brachiopods (Neobolus warthi, Lingulella)Hyolithes wynnei
Plants
10Chitarwata Formation
Paleogene Mammal fossilsPlant fossils
Datta Formation
Jurassic Plant fossilsCoal tracesFreshwater bivalves
Ghazij Group
Paleogene Plant fossilsFreshwater faunaMarginal marine fauna
Indus Formation
Cretaceous Rare plant fossilsTrace fossils
Manchhar Formation
Neogene Mammal fossilsPlant fossils
Murree Formation
Neogene Mammal bonesFreshwater molluscsPlant fossils
Ranikot Formation
Paleogene Larger foraminiferaMollusksPlant fossils
Ranikot Group
Paleogene Larger foraminiferaGastropodsPlant fossils in upper part
Toi Formation
Paleogene freshwater bivalvesgastropodsplant remains
Tredian Formation
Triassic Plant fragmentsVertebrate bones (rare)
Stromatolites & algae
9Abbottabad Formation
Cambrian StromatolitesTrace fossils
Aghil Limestone
Triassic–Jurassic ForaminiferaAlgae
Hazara Formation
Neoproterozoic Stromatolites (rare)
Laki Formation
Paleogene foraminiferagastropodsbivalvesechinoidsalgae
Nisai Limestone
Cretaceous–Paleogene Miscellanea miscellaOther larger foraminiferaAlgae
Salt Range Formation
Ediacaran StromatolitesAlgal mats
Samana Suk Formation
Jurassic CoralsBrachiopodsEchinoidsForaminiferaAlgae
Sinjrani Volcanic Group
Cretaceous AlgaeForaminiferaCorals (in limestone interbeds)
Yashkuk formation
Jurassic benthic foraminiferacalcareous algaenannofossilsostracods
Trace fossils
9Abbottabad Formation
Cambrian StromatolitesTrace fossils
Baghanwala Formation
Cambrian Trace fossilsTrilobites (rare)
Datta Formation
Jurassic Plant fossilsCoal tracesFreshwater bivalves
Hoshab Formation
Paleogene–Neogene Planktonic foraminiferaTrace fossils
Indus Formation
Cretaceous Rare plant fossilsTrace fossils
Khewra Sandstone
Cambrian Trilobite trails (trace fossils)Skolithos burrowsCruziana traces
Pab Sandstone
Cretaceous BivalvesGastropodsTrace fossils
Panjgur Formation
Paleogene–Neogene ForaminiferaTrace fossils
Siahan Formation
Paleogene Deep water foraminiferaTrace fossils
Conodonts & ostracods
11Baroghil group
Ordovician–Silurian acritarchsconodontsbrachiopods
Borom Formation
Triassic ConodontsForaminifera
Jafar Kandao Formation
Devonian–Carboniferous conodonts
Kashala formation
Triassic conodonts
Mianwali Formation
Triassic Claraia bivalvesOphiceras ammonoidsConodontsFish remains
Nowshera Formation
Devonian coralsstromatoporoidscephalopodsgastropodsbrachiopodsconodonts
Panjpir Formation
Ordovician–Silurian crinoidspelecypodscephalopodsnautiloidsconodonts
Reshit Formation
Jurassic foraminiferaostracodscrinoidscoralsechinoidsbrachiopodsammonites
Shogram Formation
Devonian rugose coralsbrachiopodsconodonts
Yashkuk formation
Jurassic benthic foraminiferacalcareous algaenannofossilsostracods
Zait limestone
Triassic conodonts
Fossil groupings are derived from each unit’s recorded fossil note and are a guide for browsing, not a formal palaeontological classification. Source: GSP Memoir Vol. 24 and the wider literature.